Publication - Factsheet

Marine science: directory of scientists

Published: 23 Nov 2019

Details of scientists working for the Marine Scotland Directorate, with some supporting documents attached.

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Marine science: directory of scientists
Dr Peter J Wright

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Dr Peter J Wright

Telephone: +44 (0)131 244 3224

E-mail: P.J.Wright@marlab.ac.uk (main address) or Peter.Wright@gov.scot

Marine Scotland Science, Marine Laboratory, 375 Victoria Road, Aberdeen, AB11 9DB, Scotland, UK

Research Interests

My key research interests are in population connectivity, fish reproductive biology, climate effects on fish populations and the application of otoliths in the study of early fish life history. Currently, my main work is in support of the Scottish MPA project and fish conservation measures.

Collaborations

I collaborate with scientists across the UK, Europe and North America, including within ICES and NAFO working groups and EU funded projects. My current collaborations are with the Universities of Aberdeen, Glasgow and Stirling, the  Danish Technical University and DFO, Canada.

External Committees and Responsibilities

UK representative ICES Science Committee, Board member Scottish MPA project (representing MSS)

Former chair of STECF and ICES  working groups on sandeel fisheries ; Cell co-ordinator European Fish Ageing Network, Member of NAFO and COST working groups on Reproductive potential, Editorial board of Journal of Fish Biology and former Council Member of Fisheries Society of the British Isles.

Most Recent Publications

  • Holmes, S., Millar, C., Fryer, R., and Wright, P.J. 2014. Gadoid dynamics: differing perceptions when contrasting stock versus population trends and its implications to management. ICES Journal of Marine Science. doi:10.1093/icesjms/fsu075
  • Neat, F., Pinto, C., Burrett, I., Cowie, L., Travis, J., Thorburn, J., Gibb, F., Wright, PJ. 2014. Site-fidelity, survival and conservation options for the threatened flapper skate (Dipturus intermedia). Aquatic Conservation Marine and Freshwater Ecosystems.DOI: 10.1002/aqc.2472
  • Wright, P.J. 2013. Methodological challenges to examining the causes of variation in stock reproductive potential. Fisheries Research, 138: 14 - 22.
  • Wright, P.J., Tobin, D 2013 Maturation differences between sub-stocks of haddock, Melanogrammus aeglefinus. Marine Biology, 160: 231-239.
  • Wright, P. 2014. Are there useful life history indicators of stock recovery rate in gadoids? ICES Journal of Marine Science. DOI:10.1093/icesjms/fsu100

For further information on research interests go to ResearchGate.

Full publications list

Anderwald, P., Evans, P., Dyer, R., Dale, A., Wright, P., and Hoelzel, A. (2012). Spatial scale and environmental determinants in minke whale habitat use and foraging. Marine Ecology Progress Series 450, 259-274.

Appelberg, M., N. Formigo, A. J. Geffen, C. Hammer, W. McCurdy, J. Modin, E. Moksness, H. Mosegaard, B. Morales-Nin, H. Troadec and P. Wright (2005). EFAN & TACADAR: A cooperative effort to exchange experience and protocols between European fish institutes. Fisheries Research 76, 167–173.

Armstrong, J.D., Fallon-Cousins, P., and Wright, P.J. (2004). The relationship between specific dynamic action and otolith growth in a teleost, northern pike. Journal of Fish Biology 64, 739-749.

Boulcott, P., Wright, P.J. (2011). Variation in fecundity in the lesser sandeel: implications for regional management. Journal of Marine Biological Association 91, 1273-1280.

Boulcott, P., Wright, P., and Gibb, F. (2007). Regional variation in maturation of sandeels in the North Sea. ICES Journal of Marine Science 64, 369–376.

Boulcott, P., and Wright, P.J. (2008). Critical timing for reproductive allocation in a capital breeder: evidence from sandeels. Aquatic Biology 3, 31-40.

Castro, C., Wright, P., Millar, C., and Holmes, S. (2013). Evidence for sub-stock dynamics within whiting (Merlangius merlangus) management regions. ICES Journal of Marine Science.

Devine, Jennifer A., Wright, Peter J., Pardoe, Heidi E., and Heino, M. (2012). Comparing rates of contemporary evolution in life-history traits for exploited fish stocks. Canadian Journal of Fisheries and Aquatic Sciences 69, 1105-1120.

Dickey-Collas, M., Armstrong, M.J., Officer, R.A., Wright, P.J., Brown, J., Dunn, M.R., and Young, E.F. (2003). Growth and expansion of haddock (Melanogrammus aeglefinus L.) stocks to the west of the British Isles in the 1990's. ICES Marine Science Symposia 219, 271-282.

Fox, C.J., Taylor, M., Dickey-Collas, M., Fossum, P., Kraus, G., Rohlf, N., Munk, P., van Damme, C.J., Bolle, L.J., Maxwell, D.L., et al. (2008). Mapping the spawning grounds of North Sea cod (Gadus morhua) by direct and indirect means. Proceedings of the Royal Society B:  Biological  Science 275, 1543-1548.

Frederiksen, M., Wright, P.J., Wanless, S., Harris, M.P., Mavor, R.A., and Heubeck, M. (2005). Regional variation in black-legged kittiwake Rissa tridctyla breeding success in Britain and Ireland and its relation to sandeel population structure. Marine Ecology Progress Series 300, 201–211.

Galley, E., Wright, P.J., and Gibb, F. (2006). Combined methods of otolith shape analysis improve identification of spawning areas of Atlantic cod. ICES Journal of Marine Science 63, 1710-1717.

Geffen, A.J., de Pontual, H., Wright, P.J., and Mosegaard, H. (2002). Chapter IIIb Life History. In Manual of Fish Sclerochronology, J. Panfili, H. Troadec, H. de Pontual, and P.J. Wright, eds. (IRD/IFREMER).

Gibb, F.M., Gibb, I.M., and Wright, P.J. (2007). Isolation of Atlantic cod (Gadus morhua) nursery areas. Marine Biology 151, 1185-1194.

Harrald, M., Neat, F.C., Wright, P.J., Fryer, R.J., and Huntingford, F.A. (2010). Population variation in thermal growth responses of juvenile Atlantic cod (Gadus morhua L.). Environmental Biology of Fishes 87, 187-194.

Harrald, M., Wright, P.J., and Neat, F.C. (2010). Substock variation in reproductive traits in North Sea cod (Gadus morhua). Canadian Journal of Fisheries and Aquatic Sciences 67, 866-876.

Heath, M.R., Culling, M.A., Crozier, W.W., Fox, C.J., Gurney, W.S.C., Hutchinson, W.F., Nielsen, E.E., O'Sullivan, M., Preedy, K.F., Righton, D.A., et al. (2013). Combination of genetics and spatial modelling highlights the sensitivity of cod (Gadus morhua) population diversity in the North Sea to distributions of fishing. ICES Journal of Marine Science: Journal du Conseil.

Heath, M.R., Kunzlik, P.A., Gallego, A., Holmes, S.J., and Wright, P.J. (2008). A model of meta-population dynamics for North Sea and West of Scotland cod—The dynamic consequences of natal fidelity. Fisheries Research 93, 92-116.

Heath, M.R., Neat, F.C., Pinnegar, J.K., Reid, D.G., Sims, D.W., and Wright, P.J. (2012). Review of climate change impacts on marine fish and shellfish around the UK and Ireland. Aquatic Conservation: Marine and Freshwater Ecosystems 22, 337-367.

Hedger, R., McKenzie, E., Heath, M., Wright, P., Scott, B., Gallego, A., and Andrews, J. (2004). Analysis of the spatial distributions of mature cod (Gadus morhua) and haddock (Melanogrammus aeglefinus) abundance in the North Sea (1980–1999) using generalised additive models. Fisheries Research 70, 17-25.

Hislop, J.R.G., Gallego, A., Heath, M.R., Kennedy, F.M., Reeves, S.A., and Wright, P.J. (2001). A synthesis of the early life history on angler fish, Lophius piscatorius L., in northern British waters. ICES Journal of Marine Science 58, 70-86.

Holmes, S., Millar, C., Fryer, R., and Wright, P. (2014). Gadoid dynamics: differing perceptions when contrasting stock versus population trends and its implications to management. ICES Journal of Marine Science.doi:10.1093/icesjms/fsu075

Holmes, S., Wright, P., and Fryer, R.J. (2008). Evidence from survey data for regional variability in cod dynamics in the North Sea and West of Scotland. ICES Journal of Marine Science 65, 206–215.

Huntingford, F.A., and Wright, P.J. (1989). How sticklebacks learn to avoid dangerous feeding patches. Behavioural Processes 19, 181-189.

Huntingford, F.A., and Wright, P.J. (1992). The development of adaptive variation in predator avoidance in freshwater fishes. Marine Behaviour and Physiology XXII, 45-61.

Huntingford, F.A., and Wright, P.J. (1992). Inherited population differences in avoidance conditioning in three-spinedsticklebacks, Gasterosteus aculeatus. Behaviour 122, 264-273.

Huntingford, F.A., Wright, P.J., and Tierney, J. (1994). The evolution of predator avoidance in the three-spined stickleback. In Evolutionary Biology of the Threespine Stickleback, M.A. Bell, and S.A. Foster, eds. (Oxford University Press).

Jensen, H., Rindorf, A., Wright, P.J., and Mosegaard, H. (2011). Inferring the location and scale of mixing between habitat areas of lesser sandeel through information from the fishery. ICES Journal of Marine Science 68, 43-51.

Jensen, H., Wright, P.J., and Munk, P. (2004). Vertical distribution of pre-settled sandeel (Ammodytes marinus) in the North Sea in relation to size and environmental variables. ICES Journal of Marine Science 60, 1342-1351.

Kalish, J.M., Beamish, R.J., Brothers, E.B., Casselman, J.M., Francis, C., Mosegaard, H., Panfili, J., Prince, E.D., Thresher, R.E., Wilson, C.A., et al. (1995). Glossary. In Recent Developments in Fish Otolith Research, D.H. Secor, J.M. Dean, and S.E. Campana, eds. (Columbia, SC, USA, Baruch Library in Marine Science, University of South Carolina Press), pp. 723-730.

Lambert, Y.N.A., Yaragina, G., Kraus, G., Marteinsdottir, G., and Wright, P.J. (2003). Correlation between reproductive characteristics and environmental and biological indices as alternative methods of estimating egg and larval production. Journal of Northwest Atlantic Fishery Science 33, 115-159.

Lewis, S., Wanless, S., Wright, P.J., Harris, J., Bull, J., and Elston, D.A. (2001). Diet and breeding performance of black-legged kittiwakes, Rissa tridactyla, at a North Sea colony. Marine Ecology Progress Series 221, 277-284.

Lynam, C., Halliday, N., Höffle, H., Wright, P., van Damme, C., Edwards, M., and Pitois, S. (2013). Spatial patterns and trends in abundance of larval sandeels in the North Sea: 1950-2005. ICES Journal of Marine Science 70 (3), 540-553.

Marshall, C.T.L., O'Brien, J., Tomkiewicz, G., Marteinsdóttir, M.J., Morgan, F., Saborido-Rey, F., Köster, F., Blanchard, D., Secor, G., Kraus, G., et al. (2004). Incorportating alternative indices of reproductive potential into fisheries management: case studies for stocks spanning an information gradient. Journal of Northwest Atlantic Fishery Science 33, 161-190.

Marshall, C.T.O.B., L. ; Tomkiewicz, Jonna ; Marteinsdottir, G. ; Morgan, M.J. ; Saborido-Rey, F. ; Köster, Fritz ; Blanchard, J.L. ; Secor, D.H. ; Kraus, Gerd ; Wright, P. ; Mukhina, N.V. ; Björnsson, H. (2003). Developing alternative indices of reproductive potential for use in fisheries management: case studies for stocks spanning an information gradient. Journal of Northwest Atlantic Fishery Science 33, 161-190.

Marubini, F., Gimona, A., Evans, P.G.H., Wright, P.J., and Pierce, G.J. (2009). Habitat preferences and interannual variability in occurrence of the harbour porpoise Phocoena phocoena off northwest Scotland. Marine Ecology Progress Series 381, 297-310.

Metcalfe, N.B., Wright, P.J., and Thorpe, J.E. (1992). Relationships between social status, otolith size at first feeding and subsequent growth in Atlantic salmon (Salmo salar ). Journal of Animal Ecology 61, 585-589.

Monaghan, P., Walton, P., Austin, G., Burns, M.D., Turner, C.M., and Wright, P.J. (1997). Sub-lethal effects of the Braer oil spill on seabirds. In: Proceedings of the Royal Society of Edinburgh Symposium on the Braer. In The Impact of An Oil Spill in Turbulent Waters: The Braer, J.M. Davies, and G. Topping, eds. (Edinburgh, The Stationary Office).

Monaghan, P., Wright, P.J., Bailey, M.C., Uttley, J.D., and Walton, P. (1996). The influence of changes in food abundance on diving and surface feeding seabirds. In Studies of high-latitude seabirds 4 Trophic relationships and energetics of endotherms in cold ocean systems, W.D. Montevecchi, ed. (Canadian Wildlife Service Occasional Paper).

Morgan, M.J., Wright, P.J., and Rideout, R.M. (2013). Effect of age and temperature on spawning time in two gadoid species. Fisheries Research 138, 42-51.

Mosegaard, H., Folkvord, A., and Wright, P.J. (2002). Ecological applications of individual data. In Manual of Fish Sclerochronology, J. Panfili, H. Troadec, H. de Pontual, and P.J. Wright, eds. (IRD/IFREMER).

Munk, P., Wright, P.J., and Pihl, N.J. (2002). Distribution of the early larval stages of cod, plaice and lesser sandeel across haline fronts in the North Sea. Estuarine, Coastal and Shelf Science 55, 139-149.

Nash, R., Wright, P., Matejusova, I., Dimitrov, S., O'Sullivan, M., Augley, J., and Hoffle, H. (2012). Spawning location of Norway pout (Trisopterus esmarkii Nilsson) in the North Sea. ICES Journal of Marine Science 69, 1338-1346.

Neat, F., Pinto, C., Burrett, I., Cowie, L., Travis, J., Thorburn, J., Gibb, F., and Wright, P. (2014). Site-fidelity, survival and conservation options for the threatened flapper skate (Dipturus intermedia). Aquatic Conservation Marine and Freshwater Ecosystems. DOI: 10.1002/aqc.2472

Neat, F., Wright, P., Zuur, A., Gibb, I., Gibb, F., Tulett, D., Righton, D., and Turner, R. (2006). Residency and depth movements of a coastal group of Atlantic cod ( Gadus morhua L.). Marine Biology 148, 643-654.

Neat, F.C., Breen, M., Cook, R.M., Gibb, I.M., and Wright, P.J. (2009). Electronic tags reveal behaviour of captured and discarded fish. Journal of Fish Biology 74, 715-721.

Neat, F.C., Wright, P.J., and Fryer, R.J. (2008). Temperature effects on otolith pattern formation in Atlantic codGadus morhua. Journal of Fish Biology 73, 2527-2541.

Neuenfeldt, S., Righton, D., Neat, F., Wright, P., Svedäng, H., Michalsen, K., Subbey, S., Steingrund, P., Thorsteinssson, V., Pampouli, C., et al. (2013). Migrations of Atlantic cod in the NE Atlantic: then, now and the future. Journal of Fish Biology 82, 741-763.

Nielsen, E.E., Wright, P.J., Hemmer-Hansen, J., Poulsen, N.A., Monro Gibb, I., and Meldrup, D. (2009). Microgeographical population structure of cod Gadus morhua in the North Sea and west of Scotland: the role of sampling loci and individuals. Marine Ecology Progress Series 376, 213-225.

Pedersen, S.A., Lewy, P., and Wright, P.J. (1999). Assessments of the lesser sandeel (Ammodytes marinus) in the North Sea based on revised stock divisions. Fisheries Research 41, 221-241.

Poloczanska, E.S., Cook, R.M., Ruxton, G.D., and Wright, P.J. (2004). Fluctuations in sandeel biomass at Shetland: implications for the commercial fishery and breeding seabirds. ICES Journal of Marine Science 61, 788-797.

Proctor, R., Wright, P.J., and Everitt, A. (1998). Modelling the transport of larval sandeels on the north west European shelf. Fisheries Oceanography 7, 347-354.

Righton, D.A., Andersen, K.H., Neat, F., Thorsteinsson, V., Steingrund, P., Svedäng, H., Michalsen, K., Hinrichsen, H.H., Bendall, V., Neuenfeldt, S., et al. (2010). Thermal niche of Atlantic cod Gadus morhua: limits, tolerance and optima. Marine Ecology Progress Series 420, 1-13.

Scott, B.E., Marteinsdottir, G., and Wright, P.J. (1999). The potential effects of maternal factors on spawning stock - recruitment relationships under varying fishing pressure. Canadian Journal of Fisheries and Aquatic Sciences 56, 1882-1890.

Scott, B.E., Marteinsdottir, G., Begg, G.A., Wright, P.J., and Kjesbu, O.S. (2006). Effects of population size/age structure, condition and temporal dynamics of spawning on reproductive output in Atlantic cod (Gadus morhua). Ecological Modelling 191, 383-415.

Swan, S., Wright, P., Woodroffe, D., Gordon, J.D.M., and Shimmield, T. (2004). Evidence for geographical isolation of the early life stages of the white anglerfish, Lophius piscatorius, based on otolith microchemistry. Journal of the Marine Biological Association of the United Kingdom 84, 827-830.

Tobin, D., and Wright, P.J. (2011). Temperature effects on female maturation in a temperate marine fish. Journal of Experimental Marine Biology and Ecology 403, 9-13.

Tobin, D., Wright, P.J., Gibb, F.M., and Gibb, I.M. (2010). The importance of life stage to population connectivity in whiting (Merlangius merlangus) from the northern European shelf. Marine Biology 157, 1063-1073.

Tobin, D., Wright, P.J., and O'Sullivan, M. (2010). Timing of the maturation transition in haddock Melanogrammus aeglefinus. Journal of Fish Biology 77, 1252-1267.

Wanless, S., PJ, W., Harris, M., and Elston, D.A. (2004). Evidence for decrease in size of lesser sandeels Ammodytes marinus in a North Sea aggregation over a 30-yr period. Marine Ecology Progress Series 279, 237–246.

Woodroffe, D.A., Wright, P.J., and Gordon, J.D.M. (2003). Verification of annual increment formation in the white anglerfish, Lophius piscatorius, using the illicia and sagitta otoliths. Fisheries Research 60, 345-356.

Wright, PJ and Gibb, F.M. (2005). Selection of birth date in North Sea haddock and its relation to maternal age. Journal of Animal Ecology 74, 303–312.

Wright, P.J., Tobin, D (2013 ). Maturation differences between sub-stocks of haddock, Melanogrammus aeglefinus. Marine Biology 160, 231-239.

Wright, P. (2014). Are there useful life history indicators of stock recovery rate in gadoids? ICES Journal of Marine Science. doi: 10.1093/icesjms/fsu100

Wright, P., Gibb, F., Gibb, I., and Millar, C. (2011). Reproductive investment in the North Sea haddock: temporal and spatial variation. Marine Ecology Progress Series 432, 149-160.

Wright, P.J. (1990). The reproductive strategy of Encrasicholina (= Stolephorus) heteroloba in the south Java Sea. Paper presented at: Tuna baitfish in the Indo-Pacific region (Honiara, Solomon Islands, ACIAR Proceedings ).

Wright, P.J. (1991). Calcium binding by soluble matrix of the otoliths of Atlantic salmon, Salmo salar L. Journal of Fish Biology 38, 625-627.

Wright, P.J. (1991). The influence of metabolic rate on otolith increment width in Atlantic salmon parr, Salmo salar L. Journal of Fish Biology 38, 929-933.

Wright, P.J. (1992). Ovarian development, spawning frequency and batch fecundity in Encrasicholina heteroloba. Journal of Fish Biology 40, 833-844.

Wright, P.J. (1993). Otolith microstructure of the lesser sandeel, Ammodytes marinus. Journal of the Marine Biological Association UK 73, 245-248.

Wright, P.J. (1996). Is there a conflict between sandeel fisheries and seabirds? A case study at Shetland. In Is there a conflict between sandeel fisheries and seabirds? A case study at Shetland, S.P.R. Greenstreet, and M.L. Tasker, eds. (Oxford, Fishing News Books, Blackwell Science), pp. 154-165.

Wright, P.J. (2007). Understanding the maturation process for field investigations of fisheries-induced evolution. Marine Ecology Progress Series 335, 279–283.

Wright, P.J. (2013). Methodological challenges to examining the causes of variation in stock reproductive potential. Fisheries Research 138, 14 - 22.

Wright, P.J., and Bailey, M.C. (1996). Timing of hatching in Ammodytes marinus from Shetland waters and its significance to early growth and survivorship. Marine Biology 126, 143-152.

Wright, P.J., and Begg, G.S. (1997). A spatial comparison of common guillemots and sandeels in Scottish waters. ICES Journal of Marine Science 54, 578-592.

Wright, P.J., Fallon-Cousins, P., and Armstrong, J.D. (2001). The relationship between otolith accretion and resting metabolic rate in juvenile Atlantic salmon during a change in temperature. Journal of Fish Biology 59, 657-666.

Wright, P.J., Galley, E., Gibb, I.M., and Neat, F.C. (2006). Fidelity of adult cod to spawning grounds in Scottish waters. Fisheries Research 77, 148-158.

Wright, P.J., and Huntingford, F.A. (1992). Daily growth increments in the otoliths of the three-spined stickleback, Gasterosteus aculeatus L. Journal of Fish Biology 41, 65-77.

Wright, P.J., and Huntingford, F.A. (1993). Agonistic interactions in juvenile sticklebacks (Gasterosteus aculeatus) in relation to local predation risk. Ethology 94, 248-256.

Wright, P.J., and Huntingford, F.A. (1993). Daily growth increments in the otoliths of the three-spined stickleback, Gasterosteus aculeatus L. Journal of Fish Biology 42, 65-77.

Wright, P.J., Jensen, H., and Tuck, I.D. (2000). The influence of sediment type on the distribution of the lesser sandeel, Ammodytes marinus. Journal of Sea Research 44, 243-256.

Wright, P.J., Metcalfe, N.B., and Thorpe, J.E. (1990). Otolith and somatic growth in Atlantic salmon parr, Salmo salar L. - evidence against coupling. Journal of Fish Biology 32, 241-250.

Wright, P.J., Millar, C.P., and Gibb, F.M. (2011). Intrastock differences in maturation schedules of Atlantic cod, Gadus morhua. ICES Journal of Marine Science: Journal du Conseil 68, 1918-1927.

Wright, P.J., Neat, F.C., Gibb, F.M., Gibb, I.M., and Thordarson, H. (2006). Evidence for metapopulation structuring in cod from the west of Scotland and North Sea. Journal of Fish Biology 69, 181-199.

Wright, P.J., Panfili, J., Meunier, F.J., Folkvord, A., and Mosegaard, H. (2002). Chapter IVa Direct validation. In Manual of Fish Sclerochronology, J. Panfili, H. Troadec, H. de Pontual, and P.J. Wright, eds. (IRD/IFREMER).

Wright, P.J., Panfili, J., and Morales-Nin, B. (2002). Chapter IIa Otoliths. In Manual of Fish Sclerochronology, J. Panfili, H. Troadec, H. de Pontual, and P.J. Wright, eds. (IRD/IFREMER).

Wright, P.J., Rowe, D., and Thorpe, J.E. (1991). Daily increments in the otoliths of Atlantic Salmon parr (Salmo salar L.) and the influence of environmental factors on their periodicity. Journal of Fish Biology 39, 103-113.

Wright, P.J., Stagg, R., and McIntosh, A. (1997). The impact of the Braer oil spill on sandeels around Shetland. In The Impact of An Oil Spill in Turbulent Waters: The Braer, J.M. Davies, and G. Topping, eds. (Edinburgh, The Stationary Office).

Wright, P.J., Talbot, C., and Thorpe, J.E. (1992). Otolith calcification in Atlantic Salmon parr (Salmo salar L.). Journal of Fish Biology 40, 779-790.

Wright, P.J., Tobin, D., Gibb, F.M., and Gibb, I.M. (2010). Assessing nursery contribution to recruitment: relevance of closed areas to haddock Melanogrammus aeglefinus. Marine Ecology Progress Series 400, 221-232.

Wright, P.J., and Trippel, E.A. (2009). Fishery-induced demographic changes in the timing of spawning: consequences for reproductive success. Fish and Fisheries 10, 283-304.

Wright, P.J., Willoughby, N.G., and Edwards, A.J. (1990). Growth, size and age composition of Stolephorus heterolobus in north central Java. In Tuna baitfish in the Indo-Pacific region (ACIAR Proceedings ).

Wright, P.J., Woodroffe, D.A., Gibb, F.M., and Gordon, J.D.M. (2002). Verification of first annulus formation in the illicia and otoliths of white anglerfish, Lophius piscatorius using otolith microstructure. ICES Journal of Marine Science 59, 587-593.

Wright, P.J., and Wright, K.M. (1991). A field study of the reproductive behaviour of spine-deficient sticklebacks from North Uist, Outer Hebrides. Scottish Naturalist 103, 3-22.

Yoneda, M., and Wright, P. (2005). Effect of temperature and food availability on reproductive investment of first-time spawning male Atlantic cod, Gadus morhua. ICES Journal of Marine Science 62, 1387-1393.

Yoneda, M., and Wright, P.J. (2004). Temporal and spatial variation in reproductive investment of Atlantic cod Gadus morhua in the northern North Sea and Scottish west coast. Marine Ecology Progress Series 276, 237-248.

Yoneda, M., and Wright, P.J. (2005). Effects of varying temperature and food availability on growth and reproduction in first-time spawning female Atlantic cod. Journal of Fish Biology 67, 1225–1241.


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